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- A superhelix is a molecular structure in which a helix is itself coiled into a helix. This is significant to both proteins and genetic material, such as overwound circular DNA. The earliest significant reference in molecular biology is from 1971, by F. B. Fuller: A geometric invariant of a space curve, the writhing number, is defined and studied. For the central curve of a twisted cord the writhing number measures the extent to which coiling of the central curve has relieved local twisting of the cord. This study originated in response to questions that arise in the study of supercoiled double-stranded DNA rings. About the writhing number, mathematician W. F. Pohl says: It is well known that the writhing number is a standard measure of the global geometry of a closed space curve. Contrary to intuition, a topological property, the linking number, arises from the geometric properties twist and writhe according to the following relationship: Lk= T + W, where Lk is the linking number, W is the writhe and T is the twist of the coil. The linking number refers to the number of times that one strand wraps around the other. In DNA this property does not change and can only be modified by specialized enzymes called topoisomerases. (en)
- 초나선(超螺旋, superhelix)은 나선(helix)이 다시 꼬여 나선 구조를 이룬 것이다. (ko)
- 超螺旋（英文：Superhelix）是一種本身就是螺旋狀的物質再次以螺旋狀纏繞成的一種特殊型態的分子結構。這種結構對於蛋白質和遺傳物質都非常重要。 (zh)
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- Twist (en)
- Writhe (en)
- CalugareanuTheorem (en)
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- Twist (en)
- Writhe (en)
- Călugăreanu Theorem (en)
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- 초나선(超螺旋, superhelix)은 나선(helix)이 다시 꼬여 나선 구조를 이룬 것이다. (ko)
- 超螺旋（英文：Superhelix）是一種本身就是螺旋狀的物質再次以螺旋狀纏繞成的一種特殊型態的分子結構。這種結構對於蛋白質和遺傳物質都非常重要。 (zh)
- A superhelix is a molecular structure in which a helix is itself coiled into a helix. This is significant to both proteins and genetic material, such as overwound circular DNA. The earliest significant reference in molecular biology is from 1971, by F. B. Fuller: About the writhing number, mathematician W. F. Pohl says: It is well known that the writhing number is a standard measure of the global geometry of a closed space curve. Contrary to intuition, a topological property, the linking number, arises from the geometric properties twist and writhe according to the following relationship: (en)
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- 초나선 (ko)
- Superhelix (en)
- 超螺旋 (zh)
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