Marine Attack Squadron 331 (VMA-331) was an attack squadron in the United States Marine Corps. The squadron, also known as the “Doodlebugs” and “Bumblebees,” was part of Marine Aircraft Group 31, 2nd Marine Aircraft Wing and was based out of Marine Corps Air Station Beaufort, South Carolina. The squadron fought in World War II and Operation Desert Storm. It was decommissioned as part of the post Cold War drawdown of the US Military on October 1, 1992.