Trilobites are used as index fossils to subdivide the Cambrian period. Assemblages of trilobites define trilobite zones. The Olenellus-zone has traditionally marked the top of the Lower Cambrian, and is followed by the zone. The last two zones of the Middle Cambrian are the Bathyuriscus–Elrathina-zone (contemporaneous with the Burgess Shale) and the subsequent -zone (starting at the base of the Drumian stage). These are overlain by the lowermost Upper Cambrian Cedaria-zone. The lower Middle Cambrian -zone (Spence Shale) is above the -zone. The -zone is upper Cambrian.
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