The 421st Rifle Division was raised in 1941 as an infantry division of the Red Army, and served briefly during the Great Patriotic War in that role. The division was formed from assets of the Odessa Military District after its namesake city came under siege by a mainly-Romanian force, and it immediately became part of the Separate Coastal Army. It was one of several improvised Soviet formations that saw service in this fighting, and its order of battle changed considerably over the course of its short existence. Prior to the fall of Odessa on October 16 what remained of the division was evacuated to the Crimea. On November 13 it was disbanded to provide replacement troops and equipment for other units defending the fortress of Sevastopol. The 421st was never reformed.
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